NAME Anatolinites chubutensis
AGE Late Paleocene.   AGE span: 58.7...55.8 mya
K&J CLASSIFICATION (2000) Fungi Imperfecti, Phragmosporae.
FIGURE(S)
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FIGURE REFERENCE Sepúlveda EG. 1980. Estudio palinológico de sedimentitas intercaladas en la "Serie Andesitica Andina", Cordon Oriental Futalaufquen, Chubut. English translation: A palynological study of microsediments intercalated in the "Andean Andesitic Series", Eastern Futalaufquen range, Chubut Province Part 1: Fungal remains; Asociación Geológica Argentina, Revista, v. 35, p. 248-272.
SPECIES, AUTHORITY A. chubutensis Elsik, Ediger & Bati 1990, p. 94.
LOCATION Andean Andesitic Series, northern Pentagonian Cordillera, Chubut Province, Argentina.
ORIG DESCRIPTION* (Sepúlveda) Diporate, three-celled fungal spores of "corrugated" oval shape, with one well-rounded, and one semi-flattened end; levigate wall, 2-3 µm thick, composed of two layers: an external one measuring 0.6 µm, and an internal one that is approximately 1.7 µm thick; two apical pores: the one on the more rounded end measuring approximately 2 µm in diameter, around which the wall bulges slightly inwards; the one on the other end also measuring about 2 µm in width, located at the end of a neck-like extension measuring 9 µm across the bottom, 6 µm across the top and 4 µm in height; the wall of this extension is translucent, and composed of one layer, 0.6 µm thick; cell partitions are opaque and similar in tone to the spore wall, which is slightly indented at the level of the partitions. Spore size: 26.5-36 x 20-23 µm (2 specimens).

EMENDED DESCRIPTION (Elsik, Ediger & Bati): Diporate, tetracellate, psilate fungal spores 20-23 x 26.5-36 µm overall. The spore axis is straight. The overall spore outline is broadly obovate, with noticeable indentations at the two distal septa, and a very marked offset to the much smaller proximal septum and cell. The two pores are both simple; the proximal pore is ca. 1.5 µm; the distal pore is ca. 2 µm, with the wall thinning from the inside of the cell into the pore. The spore wall is pigmented throughout, and the brown color approaches black, near opaque black, over the three main cells. The spore wall is ca. 0.5 µm over the proximal cell; it approaches 2-2.5 µm over the other cells. Sepúlveda (1980) described the wall over the distal cell as having an outer and inner layer, respectively 0.6 and 1.7 µm thick. The distal cell is the longest, ca. 9 µm.
COMMENTS* (Sepúlveda): Pluricellaesporites sp. 2 (pl. 2, fig. 4), bears a certain resemblance to this material, although it has only one simple pore. Also, its pigmentation is very different. Pluricellaesporites subcapsilaris Sheffy & Dilcher (1971, p. 46, pl. 14, fig. 59; pl. 16, fig. 59), is somewhat similar in form to our material, although it is lacking the neck-like extension with its pore, and its wall is much thinner, being composed of only one layer.

(Elsik, Ediger & Bati): Anatolinites chubutensis is larger than A. subcapsilaris (Sheffy & Dilcher 1971) Elsik et al. 1990. The outline of A. chubutensis also is more markedly indented over the septa.
PUBLICATION REFERENCE Sepúlveda EG. 1980. Estudio palinológico de sedimentitas intercaladas en la "Serie Andesitica Andina", Cordon Oriental Futalaufquen, Chubut. English translation: A palynological study of microsediments intercalated in the "Andean Andesitic Series", Eastern Futalaufquen range, Chubut Province Part 1: Fungal remains; Asociación Geológica Argentina, Revista, v. 35, p. 248-272.

Elsik WC, Ediger VS, Bati Z. 1990. Fossil fungal spores: Anatolinites nov. gen.; Palynology, v. 14, p. 91-103.
K&J REMARKS
TYPE Diporicellaesporites sp. 1, Sepúlveda 1980, p. 265, pl. 2, fig. 3 (stored in Argentine Mus. Natural Sci. "Bernardino Rivadavia"). Technically the name of the species A. chubutensis may not have been validly published in 1990; it is here validated by citing the information on where the type is stored (ICBN, Art. 37.5).
ALL NAMES (Including synonyms) Diporicellaesporites sp. 1, Sepúlveda 1980, p. 265, pl. 2, fig. 3 (stored in Argentine Mus. Natural Sci. "Bernardino Rivadavia").; Anatolinites chubutensis
SERIAL NUMBER 22
PUBLIC COMMENTS
 *For source, see Publication Reference.